Scrum-half Jamie Dobie, tighthead prop Murphy Walker and back-row Josh Bayliss are the other three players who have been released from the squad.
Eighteen members of the squad will be going to their first World Cup, with 23-year-old full-back Ollie Smith the youngest while new Edinburgh stand-off Ben Healy is the least capped with two.
Grant Gilchrist, Scott Cummings, Sam Skinner and Gray – whose brother Jonny was out of contention through injury – are the four locks. Russell and Healy are the two stand-off options while Blair Kinghorn, who can also deputise at 10, and Smith are the full-backs and Darcy Graham, Duhan van der Merwe and Kyle Steyn make up an exciting wing trio.
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